1983. . XIX, 1. - . 58-63 575.11:599.323.4 tII. , (t- . ), , . Mus musculus L. , >50%. t- , . musculus (t0, t9, t12 . tw5), , t. – t- t- – (tv – viable), tv 1 t [3, : (lethal – t1) [1, 2]. sl (semilethal – t ) . tv ; , 4]. tsl 2 , t v- t wPa-3 tsl , (twPa-2 , [5]. : ), tv- [4], 51% . , (tw2) tsl 51%- [6] tsl tsl . t- « » . tsl 100%- , , 10% . , », tv twPa-2 twPa-3), tsl , tsl , 10-100%. , , - . – tl . , , , , - tl . [2, 4, 7, 8]. , , . tl - . [9, 10]. , . musculus. tl1/tl2 - – t , 1 , , , -2 [11, 12]. , - , t(0,5); t- >0,5. t. musculus, [8]. t: « » « 11 , tl t- - » [13, 14]. , , tw5). . , (t0, t9, t12 , , +/tw tw – [2]). , , ; [13] ( . wild , , (T/t) T/tw. . T/tw (T/t w . +/tw /tw) . tw t v-, tsl( [ T/tw (Stw +/+ × - tw – ) Stw tl , +/tw T/tw +/tw]), 0 9 12 : t,t,t tw5 [8]. T/t . tw , ( , - tw , T/tw t/t - ) T/tw. , , . tw [2, 10]. , tl1/tl2 (n ) ) . T/t, (N) . N=2T/t. : 0,5 ( . [2]), : T/T + tl1/tl2 = T/tl1/ + T/tl2. (T/t1), , ) ( , . tl1/tl2 K=n : n ×100%. - (K) ) , , N=2T/t. St : n=N:2×St. 3 ( * 0,01%- (tw ) : «+» – 5%; «—» – ; «++» – 1%- ; «+++» – . .1 tw tw , (T/tw , /tw) (Stw) - . : (T/tw) . , tv (2,6-3,8 ) tsl. - , (3,0-3,8 (St 0,80) ). - (St 0,80) . (1:1) . t- , [2, 15]. .2 t12- t0-, t9-, (t6, twl8, t12, tw5), . , . ., tw5 , [2, 8, 9]. ), tw ( tw . . . 2), , tw K ( , . .2 3) t- [9]. , , , tw , culus - . 3). ( . , . mus[2, 4, ), , 15]. – tw5, , . , tw5 , [4, 16, 17]. . musculus , , [18], , , . – tw73 [4, 16]. twPa-l, , tw : twl ( ), tw73 twPa~l . musculus). .1 1 t -, /t t sl tw , . , tsl t /t , sl , sl sl tsl/tsl [5]. (T/tw - /tw) . tw , , , tw , tsl 10%- . tsl 2 twM45, twM08, twM010 , twM024 t1 . t1-, , [2]. twM16, twM17-6, twM41, twM28, , twM07, twM017 twM018 - ts1 . , (T/t× /t), t1-, ts1 . tw . musculus w (t ) (1:1). , - , , - , [6, 15]. tw . 3). , , , tw Kcp ( tw , . - , . , , . , , tw , ( . . 3). . , Kcp « , » « » tl . 1. Bennett D., Dunn L. . Transmission ratio distorting genes on chromosome IX and their interaction. –In: Immunogenetics of the H-2 System. Karger, Basel, 1971, p. 90. 2. Bennett D. The T-locus of the mouse. -Cell, 1975, v. 6, p. 441. 3. Braden A. W. H. T-locus in mice: segregation distortion and sterility in the male. – In: The genetics of the spermatozoon.– Proc. Int. Sympos. Edinburgh, 1972, p. 289. 4. Guenet J.L., Condamine H., Gaillard J., Jacob F. twPa-1, twPa-2, twPa-3: three new t-haplotypes in the mouse. - Genet. Res., 1980, v. 36. 2, p. 211. 5. Bennett D., Dunn L. C. Genetical and embriological comparisons of semilethal t alleles from wild mouse populations. – Genetics, 1969, v. 61, 2, p. 411. 6. Lewontin R. C. The effect of differential viability on the population dinamics of t-alleles in the mouse.- Evolution, 1968, v. 22, 2, p. 262. 7. Artzl K., Rabiarz ., Bennett D. A t-haplotype (tw35) overlapping two complementation groups. Genet. Res., 1979, v. 33, 3, p. 279. 8. . ., . . (Mus musculus L.).– .: . . 9. .: , 1980, . 97. 9. Silagi S. A genetical and embriological study of partial complementation between lethal alleles at the T locus of the house mouse. –Develop. Biol., 1962, v. 5, 1, p. 35. 10. . . (Mus musculus L.). – . ., 1980, . 14, 2, . 73. 11. Hammerberg G., Klein J., Artzt K., Bennett D. Histocompatibility-2-system in wild mice. II. H-2 haplotypes of t-bearing mice. – Transplantation. 1976, v. 21, 3, p. 199. 12. Snell G. D. The H-2 locus of the mouse: observations and speculations concerning its comparative genetics and its polymorphism. –Folia Biol. (Praha), 1968, v. 14, 5, p. 335. 13. , ., . t. I. tMus musculus L. .. 1980, . 16, 7, . 1270. 14. . , . ., . . (Mus musculus L.).– . ., 1981, . 16, 12, . 182. 15. . : Mus musculus.– .: . .: , 1977, . 39. 16. Dunn L. , Bennett D., Cookingham J. Polymorphisms for lethal alleles in European populations of Mus musculus. – J. Mammal., 1973. v. 54, 4, p. 822. 17. Klein J., Hammerberg The control of differentiation by the T complex. – Immunol. Revs, 1977, v. 33, p. 70. 18. Chaline J. Rodents, evolution and prehistory. – Endeavour, 1977, v. 1, 2, p. 44. 29.V.1981 . . , 25.IX.1984 STUDY OF t-HAPLOTYPES OF NATURAL MOUSE POPULATIONS (MUS MUSCULUS L.) II. ANALYSIS OF COMPLEMENTATION GROUPS DEMIN Yu. S., KRYUKOV V. I. A. N. Severtzov Institute of Evolutionary Morphology and Ecology of Animals, Academy of Sciences of the USSR, Moscow Summar w A description of eleven t -haplotypes (T locus, IX linkage group of Mus musculus L.) is presented which were found in two natural populations of mice from Tajikistan (USSR). These haplotypes are recessive lethals with the transmission ratio 0.73-1.00. It was shown that they do not belong to the t0, t9, t12 and tw5 complementation groups. : , , II. 1. – . 58-63 , . t// . . –1983. – . XIX,